BOOK ON CARIDEAN SHRIMPS:  Bauer, R.T. (March, 2004).  Remarkable Shrimps: Natural History and Adaptations of the Carideans. University of Okalahoma Press, Norman.  (hardcover, 316 pp., 110 figures, 11 color and half-tone plates) (ISBN-0-8061-3555-7, $59.95). Order book at University of Oklahoma Press or call 800-627-7377 or fax 800-735-0476; or try Amazon Books

Recent and Cited Publications: All publications are accessible as  pdfs at the Decapoda ATOL (a Tree of Life) website.
(; If you don't find them there, just email me for one (

Osawa,Y., M. N. Aoki, R.T. Bauer, M. Thiel, Numbers and sizes of shrimps Rhynchocinetes uritai Kubo (Decapoda: Caridea) caught in bait and refuge traps. Journal of Crustacean Biology 35: 786-775.

Osawa,Y., M. N. Aoki, M.Thiel, and R.T. Bauer. 2015. Analysis of Life-History Traits in a Sex-Changing Marine Shrimp (Decapoda: Caridea: Rhynchocinetidae). Biological Bulletin, 228: 125–136.

Rasch, J.A. and R.T. Bauer. 2015. Temporal variation in population structure of the isopod Urobopyrus processae Richardson, 1904 (Isopoda: Bopyridae) infesting the branchial chamber of the night shrimp Ambidexter symmetricus Manning and Chace, 1971 (Decapoda: Processidae). Nauplius 23: 89-103.

Rasch, J.A. and R.T. Bauer. 2015. Reproductive pattern and sexual system of the nocturnal seagrass shrimp Ambidexter symmetricus (Decapoda: Caridea: Processidae) in a Florida bay. Marine & Freshwater Research.

Sanjeevi P., T. T. Ajithkumar, R. Bauer, M. Thiel, and T. Subramoniam. 2015. Reproductive morphology and mating behavior in the hingebeak shrimp Rhynchocinetes durbanensis Gordon, 1936 (Decapoda: Caridea: Rhynchocinetidae) in India. Journal of the Marine Biological Society, U.K. doi:10.1017/S0025315415001083

Bauer, R.T. 2015. Review: Atlas of Crustacean Larvae. J.W. Martin, J. Olesen, and J.T. Høeg. Johns Hopkins University Press, Baltimore, 370 pp.. Quarterly Review Biology 90:217-218.

Solomon T. C. Chak, Raymond Bauer, Martin Thiel. 2015. Social Behaviour and Recognition in Decapod Shrimps, with Emphasison the Caridea . L. Aquiloni and E. Tricarico (eds.), Social Recognition in Invertebrates, pp.57-84. Springer International Publishing, Cham, Switzerland.

Castilho, A.L., R. T. Bauer; F.A. M. Freire; V. Fransozo; R. C. Costa; R. C. Grabowski and A. Fransozo. 2015. Lifespan and reproductive dynamics of the commercially important sea bob shrimp Xiphopenaeus kroyeri (Penaeoidea): synthesis of a 5-year study. Journal of Crustacean Biology 35: 30-40.

Bauer, R., Gimenez , and V. Storch. 2014. Klaus Anger, Recipient of The Crustacean Society Excellence In Research Award. Journal of Crustacean Biology 35: 116-119.

Bauer, R.T., J. Okuno, and M. Thiel. 2014. Inferences on mating and sexual systems of two Pacific Cinetorhynchus shrimp s (Decapoda, Rhynchocinetidae) based on sexual dimorphism in body size and cheliped weaponry. Zookeys 457: 187-209.

Bauer, R.T. 2014. Review: Jensen, Gregory C. 2014. Crabs and Shrimps of the Pacific Coast. A Guide to Shallow-water Decapods from Southeastern Alaska to the Mexican Border. Journal of Crustacean Biology 34: 532 –533 .

Baeza, J.A., Bauer, R.T., Okuno, J. and M. Thiel 2014.Molecular phylogeny of hinge-beak shrimps (Decapoda: Caridea: Rhynchocinetes and Cinetorhynchus ) and allies: a formal test of familiar and generic monophyly using a multilocus phylogeny. Zoological Journal of the Linnean Society.172 426–450 .

Gregati, R.A., Fransozo, V., López-Greco, L.S., Negreiros-Fransozo, M.L. and R.T. Bauer. 2014. Functional morphology of the reproductive system and sperm transfer in Stenopus hispidus (Crustacea: Decapoda: Stenopodidea) and their relation to mating system. Invertebrate Biology 133: 381–393

Bauer, R.T. 2013. Review: Schram, F. R., and J. C. von Vaupel Klein (eds) 2012. Treatise on Zoology–Anatomy, Taxonomy, Biology. The Crustacea. Volume 9, Part B. Eucarida: Decapoda: Astacidea P.P. (Enoplometopoidea, Nephropoidea),Glypheidea, Axiidea, Gebiidea, and Anomura.Decapoda. Journal of Crustacean Biology 33: 445-448.

Bauer, R.T. 2013. Adaptive modification of appendages for grooming (cleaning; antifouling) and reproduction in the Crustacea, pp. 337-375. In: M. Thiel and L. Watling (eds.), Functional Morphology of Crustacea, Vol. 1.  Oxford University Press, New York.

Bauer, R.T. 2013. Amphidromy in shrimps: a life history pattern between rivers the sea. Latin American Journal of Aquatic Research 41: 633-650

T. J. Olivier, S.L. Conner, and R.T. Bauer. 2012. Evidence of extended marine planktonic larval development in far-upstream populations of the river shrimp Macrobrachium ohione  (Smith, 1874) from the Mississippi River. Journal of Crustacean Biology 32: 899-905
Bauer, R.T. and M. Thiel. 2011. First Description of a pure-search mating system and protandry in the shrimp Rhynchocinetes uritai (Decapoda, Caridea). Journal of Crustacean Biology 31: 286-295.

Bauer, R.T. and S.L. Conner. 2011. Gonochoric sexual system in the caridean shrimps Processa riveroi and P. bermudensis (Decapoda: Processidae) inhabiting a tropical seagrass meadows. Journal of the Marine Biological Association United Kingdom 92: 521-529

Olivier, T.J. and R.T. Bauer. 2011.  Downstream female hatching migration of the river shrimp Macrobrachium ohione in the lower Mississippi River system. American Midland Naturalist 166:379–393

Bauer, R.T. 2011a. Amphidromy and migrations of freshwater shrimps. I. Costs, benefits, evolutionary origins, and an unusual case of amphidromy.  In : A. Asakura (ed.),  New Frontiers in Crustacean Biology. Proceedings of the TCS Summer Meeting, Tokyo, 20-24 September 2009, pp. 145-156.  Brill : Leiden, The Netherlands.

Bauer, R.T. 2011b.  Amphidromy and migrations of freshwater shrimps. II. Delivery of hatching larvae to the sea, return juvenile upstream migration, and human impacts. In : A. Asakura (ed.),  New Frontiers in Crustacean Biology. Proceedings of the TCS Summer Meeting, Tokyo, 20-24 September 2009, pp. 157-168.  Brill : Leiden, The Netherlands.

Bauer, R.T.  2011. Chemical communication in decapod shrimps: the influence of mating and social systems on the relative importance of olfactory and contact pheromones.  In: T. Breithaupt and M. Thiel (eds.), Chemical Communication in Crustaceans, pp. 277-296. Springer Verlag, New York.

Bauer, R.T. 2010.  Isabel Perez Farfante: Obituary.  Journal of Crustacean Biology 30: 345-349.

Conner, S.L. and R.T. Bauer. 2010. Infection of adult migratory river shrimps, Macrobrachium ohione, by a branchial bopyrid isopod, Probopyrus pandalicola. Invertebrate Biology 129: 344-352.

Rome, N., S.L. Conner, and R.T. Bauer.  2009.  Delivery of hatching larvae to estuaries by anamphidromous river shrimp: tests of hypotheses based on larval moulting and distribution.  Freshwater Biology 54: 1924-1932.

Caskey, J.L., K.H. Hasenstein, and R.T. Bauer.  2009. Studies on contact sex pheromones of the caridean shrimp Palaemonetes pugio: I. Cuticular hydrocarbons associated withmate recognition.  Invertebrate Reproduction and Development 53: 93-103.

Caskey, J.L., G. M. Watson, and R.T. Bauer.  2009. Studies on contact sex pheromones of the caridean shrimp Palaemonetes pugio.II. The role of glucosamine in mate recognition.  Invertebrate Reproduction and Development 53: 105-116.

Baeza, J.A., C.D. Schubart, P. Zillner, S. Fuentes, and R.T. Bauer.  2009  Molecular phylogeny of shrimps from the genus Lysmata (Caridea: Lysmatidae): the evolutionary origins of protandric simulataneous hermaphroditism and social monogamy.  Biological Journal of the Linnean Society. 96, 415–424.

Bauer, R.T. and J. Delahoussaye.2008.  Life history migrations of the amphidromous river shrimp Macrobrachium ohione from a continental large river system.  Journal of Crustacean Biology 28: 622-632.

Tóth, E. and R.T. Bauer.  2008.  Synalpheus paraneptunus (Crustacea: Decapoda: Caridea) populations with intersex gonopores: a sexual enigma amongsponge-dwelling snapping shrimps.  Invert. Reprod. Develop. 51: 49-59.

Bauer, R.T.  2007.  Hermaphroditism in caridean shrimps: mating systems, sociobiology, and evolution, with special reference to Lysmata.  In: J.E. Duffy and M. Thiel.  2007.  Evolutionary ecology of social and sexual systems: crustaceans as model organisms, pp. 232—248.  Oxford University Press, New York
Toth, E. and R.T. Bauer.  2007.  Gonopore sexing technique allows determination of sex ratios and helper composition in eusocial shrimps.  Marine Biology 151: 1875-1886.

McLaughlin, P.A., R. Lemaitre, F.  D. Ferrari, D.L. Felder, and R.T. Bauer.  2008.  A Reply to T.W. Flegel.  Aquaculture 275: 370-373.

Castilho, A.L., Gavio, M.A., Costa, R.C., Boschi, E.E., Bauer, R.T., and Fransozo, A.  2007.  Latitudinal variation in population structure and reproductive pattern of the endemic South American shrimp Artemesia longinaris (Decapoda: Penaeoidea).  Journal of Crustacean Biology 27: 548-552.

Bauer, R.T. 2006.  Same sexual system but variable sociobiology: evolution of protandric simultaneous hermaphroditism in Lysmata shrimps.  Integrative and Comparative Biology 46: 430-438.

Bauer, R.T. and J.L. Caskey. 2006.  Flagellar setae of the second antennae in decapod shrimps: sexual dimorphism and possible role in detection of contact sex pheromones.  Invertebrate Reproduction and Development 49:  51-60.

Caskey, J.L. and R.T. Bauer.  2005.  Behavioral tests for a possible contact sex pheromone in the caridean shrimp Palaemonetes pugio.  Journal of Crustacean Biology 25: 35-40.

Bauer, R.T.  2005.  Costs of maleness on brood production in the shrimp Lysmata wurdemanni  (Decapoda: Hippolytidae: Caridea), a protandric simultaneous hermaphrodite.  Journal of the Marine Biological Society, United Kingdom 85: 101-106.

Bauer, R.T., and W.A. Newman. 2004. Protandric simultaneous hermaphroditism in the marine shrimp Lysmata californica (Caridea: Hippolytidae). Journal of Crustacean Biology 24: 131-139.

Baeza, J.A, and R.T. Bauer, R.T. 2004. Experimental test of socially mediated sex change in a protandric simultaneous hermaphrodite, the marine shrimp Lysmata wurdemanni (Caridea: Hippolytidae). Behavioral Ecology and Sociobiology 5: 544-550.

Baldwin, A.P. and R.T. Bauer.  2003.  Growth, survivorship, life span, and sex change in the hermaphroditic shrimp Lysmata wurdemanni (Decapoda: Caridea: Hippolytidae). Mar.  Biol. 143: 157-166.

Bauer R.T.  2002. Test of hypotheses on the adaptive value of an extended male phase in the hermaphroditic shrimp Lysmata wurdemanni (Caridea: Hippolytidae). Biol Bull 203:347–357.

Bauer, R.T. (2002).  Reproductive ecology of a protandric simultaneous hermaphrodite, the shrimp Lysmata wurdemanni (Decapoda: Caridea: Hippolytidae). Journal of Crustacean Biology.  22: 742-749
Bauer, R.T. 2002. The ineffectiveness of grooming in prevention of body fouling in the red swamp crawfish Procambarus clarkii..  Aquaculture 208: 39-49.
Hess, G.S. and R.T. Bauer. (2002).  Spermatophore transfer in the hermit crab Clibanarius vittatus (Crustacea, Anomura, Diogenidae). Journal of Morphology 253: 166-175.

Bauer, R.T.  2001. Hermafroditismo en camarones: el sistema sexual y su relación con atributos socioecológicos. Interciencia 26: 434-439.

 Bauer, R.T. and J.A. Abdalla.  2001.  Male mating tactics in the shrimp Palaemonetes pugio (Decapoda, Caridea): precopulatory mate guarding vs. pure searching.  Ethology 107: 185-199.

 Bauer, R.T. and J.A. Abdalla.  2000.  Patterns of brood production in the grass shrimp Palaemonetes pugio (Decapoda: Caridea).  Invertebrate Reproduction and Development 38: 107-113.

Bauer, R.T.  2000.  Simultaneous hermaphroditism in caridean shrimps: a unique and puzzling sexual system in the Decapoda.  Journal of Crustacean Biology 20 (Special No. 2): 116-128.

 Bauer, R.T.  1999.  Gill-cleaning mechanisms of a dendrobranchiate shrimp, Rimapenaeus  similis (Decapoda: Penaeidae): description and experimental testing of function.   Journal of Morphology 242: 125-139.

Bauer, R.T. and G.J. Holt.  1998.  Simultaneous hermaphroditism in the marine shrimp Lysmata wurdemanni (Caridea: Hippolytidae): an undescribed sexual system in the decapod Crustacea.  Marine Biology 132: 223-235.

 Bauer, R.T.  1998.  Gill-cleaning mechanisms of the crayfish Procambarus clarkii (Astacidea: Cambaridae): experimental testing of setobranch function.  Invertebrate Biology 177: 129-143.

 Bauer, R.T.  1998.  Review of: Penaeoid and Sergestoid Shrimps and Prawns of the World.  Keys and Diagnoses for the Families and Genera by Isabel Pérez Farfante and Brian Kensley.  Bulletin of Marine Science 62: 321-323.

 Bruce, A.J. and R.T. Bauer.  1997.  Nippontonia minirostris, a new genus and species of pontoniine shrimp from Okinama.  Journal of Natural History 31: 1709-1721.

 Bauer, R.T.  1996.  Role of the petasma and appendices masculinae during copulation and insemination in the penaeoid shrimp, Sicyonia dorsalis (Crustacea: Decapoda: Dendrobranchiata).  Invertebrate Reproduction and Development 29: 173-184.

 Bauer, R.T.  1996.  A test of hypotheses on male mating systems and female molting in decapod shrimp, using Sicyonia dorsalis (Decapoda: Penaeaoidea).  Journal of Crustacean Research 16: 429-437.

 Bauer, R.T. and R. VanHoy.  1996.  Variation in sexual systems (protandry, gonochorism) and reproductive biology among three species of the shrimp genus Thor (Decopoda: Caridea).  Bulletin of Marine Science 59: 53-73.

 Bauer, R.T.  1996.  Book Review: Forest, Jacques, ed. 1994.  Traité de zoologie.  Tome VII. Crustacés.  Fascicule I.  Morphologie, physiologie, reproduction, embryologie. Journal of Crustacean Biology 16: 216-218.

 Bauer, R.T. and Junmin Lin.  1994.  Temporal patterns of reproduction and recruitment in the marine shrimps Trachypenaeus similis (Smith) and T. constrictus (Stimpson) (Crustacea: Decapoda) from the Northcentral Gulf of Mexico.  Journ. Exp. Mar. Biol. Ecol. 182: 205-222.

 Bauer, R.T. 1994.  Usage of the terms thelycum and spermatheca in the repoductive morphology of the Decapoda, with special reference to the Penaeoidea.  Journal of Crustacean Biology 14: 715-721.

 Bauer, R.T. and Lin Jun Min.  1993.  Spermatophores and plug substance of the marine shrimp Trachypenaeus similis (Crustacea: Decapoda: Penaeidae): formation in the male reproductive tract and disposition in the inseminated female.  Biological Bulletin 185: 174-185.

Cash, C.E. & R.T. Bauer.  1993.  Adaptations of the branchial parasite Probopyrus pandalicola (Isopoda: Bopyridae) for survival and reproduction related to ecdysis of the host, Palaemonetes pugio (Caridea: Palaemonidae). Journal of Crustacean Biology 13: 111-124.

 Bauer, R.T.  1992a.  Testing generalizations about latitudinal variation in reproductive and recruitment patterns with caridean and sicyoniid shrimps. Journal of Invertebrate Reproduction & Development 22: 193-202.

 Bauer, R.T. and L. Rivera Vega.  1992b.  Pattern of reproduction and recruitment in two sicyoniid shrimp species (Decapoda, Penaeoidea) from a tropical seagrass habitat.  Journal of Experimental Marine Biology and Ecology 161:223-240.

 Bauer, R.T.  1992.  Repetitive copulation and variable success of insemination in the marine shrimp Sicyonia dorsalis (Decapoda, Penaeoidea). Journal of Crustacean Biology, 12:153-160.

 Bauer, R.T. and C.E. Cash.  1991.  Spermatophore structure and anatomy of the ejaculatory duct in Penaeus setiferus, P. aztecus and P. duorarum (Crustacea: Decapoda); homologies and functional significance.  Transactions of the American Microscopical Society, 110: 144-162.

 Bauer, R.T.  1991. Analysis of embryo production in a caridean shrimp guild inhabiting tropical seagrass meadows.  In A. Wenner & A. Kuris (eds.) "Crustacean Egg Production," Crustacean Issues, Balkema Press, 7:181-192.

 Bauer, R.T.  1991.  Sperm transfer and storage structures in penaeoid shrimps: a functional and phylogenetic perspective.  In: R.T. Bauer & J.W. Martin (eds), "Crustacean Sexual Biology," Columbia University Press, pp. 183-207.

Bauer, R.T. and J.W. Martin (eds.). 1991. Crustacean Sexual Biology, Columbia Univ. Press, 355 pp.

Bauer, R.T.  1989a.  Continuous reproduction and episodic recruitment in nine caridean shrimp species inhabiting a tropical seagrass meadow.  Journal of Experimental Marine Biology and Ecology, 127: 175-187.

 Bauer, R.T.  1989b.  Functional morphology, adaptive value, and phylogenetic significance of grooming in Decapod Crustacea.  In: B. Felgenhauer and L. Watling (eds), Functional Morphology of Grooming and Feeding Appendages, pp.49-73, Crustacean Issues, Balkema press.

 Bauer, R.T.  1989c.  Observations and experiments on grooming behavior of the tropical stomatopod Gonodactylus oerstedii.  In: Biology of Stomatopods.  E.A. Ferrero (ed.). Selected Symposia and Monographs U.Z.I., 3, Mucchi, Modena, 1989, pp. 99-104.

Bauer, R.T.  1987. "Crustacea" by F.R. Schram.  Book review in Bioscience 37: 752-754.

Bauer, R.T.  1987.  Stomatopod grooming behavior:  functional morphology and amputation experiments in Gonodactylus oerstedii.  Journal of Crustacean Biology 7: 414-432.

Bauer, R.T.  1986.  Phylogenetic trends in sperm transfer and storage complexity in decapod crustaceans.  Journal of Crustacean Biology 6: 313-325.

Bauer, R.T.  1986.  Sex change and life history pattern in the shrimp Thor manningi (Decopoda: Caridea): a novel case of partial protandric hermaphroditism. Biological Bulletin 170: 11-31.

Bauer, R.T.  1985.  Diel and seasonal variation in species composition and abundance of the caridean shrimps (Crustacea, Decapoda) from seagrass meadows in northern Puerto Rico.  Bulletin of Marine Science 36: 150-162.

Bauer, R.T.  1985.  Penaeoid shrimp fauna from tropical seagrass meadows:  species composition, diurnal and seasonal variation in abundance.  Proceedings of the Biological Society of Washington 98: 177-190.

Bauer, R.T.  1985.  Hermit crab fauna from tropical seagrass meadows: species composition, diel and seasonal variation in abundance.  Journal of Crustacean Biology 52: 249-257.

Bauer, R.T.  1984.  Morphological trends in the genus Heptacarpus (Decapoda, Caridea) and their phylogenetic significance.  Journal of Crustacean Biology 4: 201-225.

Bauer, R.T.  1982.  Polymorphism of colour pattern in the caridean shrimps  Heptacarpus pictus and H. paludicola.  Marine Behaviour and Physiology 8: 249- 265.

Bauer, R.T.  1981.  Color patterns of the shrimps Heptacarpus pictus and H. paludicola.  Marine Biology 46: 141-152.

Bauer, R.T.  1981.  Grooming behavior and morphology of the Decapod Crustacea.  Journal of Crustacean Biology 1: 153-173.

Bauer, R.T.  1979.  Antifouling adaptations of marine shrimp (Decapoda: Caridea):  gill cleaning mechanisms and grooming of brooded embryos.  Zoological Journal of the Linnean Society 65: 281-303.

Bauer, R.T.  1979. Sex attraction and recognition in the shrimp Heptacarpus  paludicola Holmes.  Marine Behaviour and Physiology 7: 157-174.

Bauer, R.T.  1978.  Antifouling adaptations of marine shrimps (Decapoda: Caridea): functional morphology and adaptive value of general body grooming and antennal cleaning behaviours.  Marine Biology 49: 69-82.

Bauer, R.T.  1977.  Antifouling adaptations of marine shrimp (Decapoda: Caridea):  functional morphology and adaptive significance of antennular preening by the third maxillipeds.  Marine Biology 40: 260-276.

Bauer, R.T.  1976.  Mating behaviour and spermatophore transfer in the shrimp Heptacarpus pictus (Stimpson) (Decapoda: Caridea: Hippolytidae).  Journal of Natural History 10: 415-440.

Bauer, R.T.  1975.  Grooming behaviour and morphology of the caridean shrimps Pandalus danae.  Zoological Journal Linnean Society 56: 45 71.
 Back to home page