Scott C. France - Publications - Return to homepage

2019 Kennedy BR, Cantwell K, Malik M, Kelley C, Potter J, Elliott K, Lobecker E, McKenna Gray L, Sowers D, France S, Auscavitch S, Mah C, Moriwake V, Bingo SRD, Putts M, Rotjan RD. The unknown and the unexplored: insights into the Pacific deep-sea following NOAA CAPSTONE expeditions. Frontiers in Marine Science 6: 480 [DOI:10.3389/fmars.2019.00480]

2017 Heestand Saucier E, Sajjadi A, France SC. A taxonomic review of the genus Acanella (Cnidaria: Octocorallia: Isididae) in the North Atlantic Ocean, with descriptions of two new species. Zootaxa 4323(3):359–390 []

2017 Wicksten M, De Grave S, France S, Kelley C. Filter-feeding in a Deep-Sea Benthic Shrimp (Decapoda: Caridea: Stylodactylidae), with Records of the Deepest Carideans. ZooKeys 646: 17-23 [doi: 10.3897/zookeys.646.10969]

2016 Lawler SN, Kellogg CA, France SC, Clostio RW, Brooke SD, Ross SW. Coral-Associated Bacterial Diversity is Conserved Across Two Deep-Sea Anthothela Species. Frontiers in Microbiology 7 [doi:10.3389/fmicb.2016.00458]

2015 Quattrini A, Nizinski MS, Chaytor JD, Demopoulos AWJ, Roark EB, France SC, Moore JA, Hey T, Auster PJ, Kinlan B, Ruppel C, Elliott KP, Kennedy BRC, Lobecker E, Skarke A, Shank TM. Exploration of the canyon-incised continental margin of the northeastern United States reveals dynamic habitats and diverse communities. PLoS ONE 10: e0139904 [doi:10.1371/journal.pone.0139904]

2015 Zapata F, Goetz FE, Smith SA, Howison M, Siebert S, Church SH, Sanders SM, Ames CL, McFadden CS, France SC, Daly M, Collins AG, Haddock SHD, Dunn CW, Cartwright P. Phylogenomic analyses support traditional relationships within Cnidaria. PLoS ONE 10: e0139068 [doi:10.1371/journal.pone.0139068]

2015 Pante E, SC France, D Gey, C Cruaud & S Samadi. An inter-ocean comparison of coral endemism on seamounts: the case of Chrysogorgia. Journal of Biogeography 42: 1907–1918 [doi:10.1111/jbi.12564

2015 Pante E, J Abdelkrim, A Viricel, D Gey, SC France, MC Boisselier & S Samadi. Use of RAD sequencing for delimiting species. Heredity 114: 450–459 (doi:10.1038/hdy.2014.105)

2015 de Moura Neves B, E Edinger, C Hillaire-Marcel, E Heestand Saucier, SC France, M Treble & V Wareham. Dense forests of a deep-water bamboo coral in a muddy Arctic environment. Marine Biodiversity 45: 867-871 (doi: 10.1007/s12526-014-0291-7)

2013 Brugler, M.R., D.M. Opresko, S.C. France. The evolutionary history of the order Antipatharia (Cnidaria: Anthozoa: Hexacorallia) as inferred from mitochondrial and nuclear DNA: Implications for black coral taxonomy and systematics. Zoological Journal of the Linnean Society 169:312-361 [ABSTRACT][doi:]

2013 Pante E., E. Heestand-Saucier, S.C. France. Molecular and morphological data support reclassification of the octocoral genus Isidoides. Invertebrate Systematics 27:365-378 [ABSTRACT][doi:]

2012 Pante E., S.C. France, A. Couloux, C. Cruaud, C.S. McFadden, S. Samadi & L. Watling. Deep-sea origin and in-situ diversication of chrysogorgiid octocorals. PLoS ONE 7: e38357 [14 pp] [ABSTRACT](Open Access: doi:10.1371/journal.pone.0038357)

2011 Watling, L., & S.C. France. A New Genus and Species of Bamboo Coral (Octocorallia: Isididae: Keratoisidinae) from the New England Seamounts. Bulletin of the Peabody Museum of Natural History 52:209-220 [ABSTRACT](doi:

2011 Watling, L., S.C. France, E. Pante & A. Simpson. Biology of Deep-Water Octocorals. Advances in Marine Biology 60:41-122 [ABSTRACT](doi: 10.1016/B978-0-12-385529-9.00002-0)

2011 de Buron, I., S.C. France, V.A. Connors, W.A. Roumillat, & L.C. Tsoi. Philometrids of the southern flounder Paralichthys lethostigma: a multidimensional approach to determine their diversity. Journal of Parasitology 97: 466-475 [ABSTRACT] (

2011 McFadden, C.S., Y. Benayahu, E. Pante, J.N. Thoma, P.A. Nevarez & S.C. France. Limitations of Mitochondrial Gene Barcoding in Octocorallia. Molecular Ecology Resources 11: 19-31 [ABSTRACT](

2010 Pante E & France SC. Pseudochrysogorgia bellona n. gen., n. sp.: a new genus and species of chrysogorgiid octocoral (Coelenterata, Anthozoa) from the Coral Sea. Zoosystema 32: 595-612 [ABSTRACT][PDF]

2010 France, S.C., E. Pante, M. R. Brugler & J.L. van der Ham. On the evolution of deep-sea octocorals and antipatharians: Patterns revealed from molecular phylogenies. Integrative and Comparative Biology 50(Suppl. 1): E56-E56 [Conference abstract]

2010 McFadden, C.S., J.A. Sanchez & S.C. France. Molecular Phylogenetic Insights into the Evolution of Octocorallia: A Review. Integrative and Comparative Biology 50(3): 389-410 [ABSTRACT](doi: 10.1093/icb/icq056)

2010 Wagner D, Brugler MR, Opresko DM, France SC, Montgomery AD & Toonen R. Using morphometrics, in situ observations and genetic characters to distinguish amongst commercially valuable Hawaiian black coral species; a redescription of Antipathes grandis Verrill, 1928 (Antipatharia: Antipathidae). Invertebrate Systematics 24: 271-290 [ABSTRACT](doi:10.1071/IS10004)

2009 Thoma, J.N., E. Pante, M.R. Brugler & S.C. France. Deep-sea octocorals and antipatharians show no evidence of seamount-scale endemism in the NW Atlantic. Marine Ecology Progress Series 397: 25-35 [ABSTRACT] (Open Access:

2009 van der Ham, J.L., M.R. Brugler & S.C. France. Exploring the utility of an indel-rich, mitochondrial intergenic region as a molecular barcode for bamboo corals (Octocorallia: Isididae). Marine Genomics 2: 183-192 [ABSTRACT] (

2008 Brugler, M.R. & S.C. France. The mitochondrial genome of a deep-sea bamboo coral (Cnidaria, Anthozoa, Octocorallia, Isididae): genome structure and putative origins of replication are not conserved among octocorals. Journal of Molecular Evolution 67: 125-136 [ABSTRACT] (

2007 Daly, M., M.R. Brugler, P. Cartwright, A.G. Collins, M.N. Dawson, S.C. France, C.S. McFadden, D.M. Opresko, E. Rodriguez, S. Romano & J. Stake. The phylum Cnidaria: A review of phylogenetic patterns and diversity 300 years after Linnaeus. In: Zhang, Z.-Q. & Shear, W.A. (Eds) (2007) Linnaeus Tercentenary: Progress in Invertebrate Taxonomy. Zootaxa 1668: 127-182 (open access pdf)

2007 France, S.C. Genetic analysis of bamboo corals (Cnidaria: Octocorallia: Isididae): does lack of colony branching distinguish Lepidisis from Keratoisis? Bulletin of Marine Science 81: 323-333 [ABSTRACT] (low res pdf 600K) (high res pdf 7.8 MB)

2007 Brugler, M.R. & S.C. France. The complete mitochondrial genome of the black coral Chrysopathes formosa (Cnidaria:Anthozoa:Antipatharia) supports classification of antipatharians within the subclass Hexacorallia. Molecular Phylogenetics and Evolution 42: 776-788 [ABSTRACT] (

2006 McFadden, C.S., S.C. France, J.A. Sánchez & P. Alderslade. A molecular phylogenetic analysis of the Octocorallia (Cnidaria: Anthozoa) based on mitochondrial protein-coding sequences (ND2, msh1). Molecular Phylogenetics and Evolution 41: 513-527 [ABSTRACT] (

2004 Smith, P. J. , S. M. McVeagh, J. T. Mingoia & S. C. France. Mitochondrial DNA sequence variation in deep-sea bamboo coral (Keratoisidinae) species in the southwest and northwest Pacific Ocean. Marine Biology 144: 253-261 [ABSTRACT] (

2003 Sánchez, J. A., C. S. McFadden, S. C. France & H. R. Lasker. Molecular phylogenetic analyses of shallow-water Caribbean octocorals. Marine Biology 142: 975-987 (

2002 France, S. C. & L. L. Hoover. DNA sequences of the mitochondrial COI gene have low levels of divergence among deep-sea octocorals (Cnidaria: Anthozoa). Hydrobiologia 471: 149-155 [ABSTRACT] (

2001 France, S. C. & L. L. Hoover. Analysis of variation in mitochondrial DNA sequences (ND3, ND4L, MSH) among Octocorallia (=Alcyonaria) (Cnidaria: Anthozoa). Bulletin of the Biological Society of Washington 10:110-118 [ABSTRACT]

2001 Berntson, E. A. & S. C. France. Generating DNA sequence information from museum collections of octocoral specimens (Phylum Cnidaria: Class Anthozoa). Bulletin of the Biological Society of Washington 10:119-129

2001 Berntson, E. A., F. M. Bayer, A. G. Mcarthur & S. C. France. Phylogenetic relationships within the Octocorallia (Cnidaria: Anthozoa) based on nuclear 18S rRNA sequences. Marine Biology 138:235-246. [ABSTRACT] (

1999 Rosel, P.E.., S. C. France, J.Y. Wang & T. D. Kocher. Genetic structure of harbour porpoise Phocoena phocoena populations in the Northwest Atlantic based on mitochondrial and nuclear markers. Molecular Ecology 8: S41-S54 [ABSTRACT] (

1999 Berntson, E. A., S. C. France & L. S. Mullineaux. Phylogenetic Relationships within the Class Anthozoa (Phylum Cnidaria) Based on Nuclear 18S rDNA Sequence Information. Molecular Phylogenetics and Evolution 13(2): 417-433 [ABSTRACT] (

1999 France, S.C., N. Tachino, T.F. Duda, Jr., R.A. Shleser & S.R. Palumbi. Intraspecific genetic diversity in the marine shrimp Penaeus vannamei: multiple polymorphic elongation factor-1alpha loci revealed by intron sequencing. Marine Biotechnology 1: 261-268 [ABSTRACT] (

1996 France, S. C. & T. D. Kocher. DNA sequencing of formalin-fixed crustaceans from archival research collections. Molecular Marine Biology and Biotechnology 5 (4): 304-313[ABSTRACT]

1996 France, S. C. & T. D. Kocher. Geographic and bathymetric patterns of mitochondrial 16S rRNA sequence divergence among deep-sea amphipods, Eurythenes gryllus.Marine Biology 126(4): 633-644 [ABSTRACT] (

1996 France, S. C., P. E. Rosel, J. E. Agenbroad, L.S. Mullineaux, & T. D. Kocher. DNA sequence variation of mitochondrial large-subunit rRNA provides support for a two subclass organization of the Anthozoa (Cnidaria). Molecular Marine Biology and Biotechnology 5 (1): 15-28 [ABSTRACT]

Mullineaux, L. S. & S. C. France. Dispersal mechanisms of deep-sea hydrothermal vent fauna. In: Seafloor Hydrothermal Systems: Physical, Chemical, Biological and Geological Interactions, S. E. Humphris, R. A. Zierenberg, L. S. Mullineaux, and R. E. Thomson, eds. Geophysical Monograph 91, American Geophysical Union, pp. 408 - 424

Brusca, R. C., R. Wetzer & S. C. France. 1995. Cirolanidae (Crustacea: Isopoda: Flabellifera) of the Tropical Eastern Pacific. Proceedings of the San Diego Society of Natural History No. 30: 1 - 96

France, S. C. 1994. Genetic population structure and gene flow among deep-sea amphipods, Abyssorchomene spp., from six California Continental Borderland basins. Marine Biology 118: 67-77 [ABSTRACT] (

Martin, J. W., S. C. France & C. L. Van Dover. 1993. Halice hesmonectes, a new species of pardaliscid amphipod (Crustacea, Peracarida) from hydrothermal vents in the eastern Pacific. Canadian Journal of Zoology 71: 1724-1732

France, S. C. 1993. Geographic variation among three isolated populations of the hadal amphipod Hirondellea gigas (Crustacea: Amphipoda: Lysianassoidea). Marine Ecology Progress Series 92:277-287 [ABSTRACT]

Van Dover, C.L., S. Kaartvedt, S.M. Bollens, P.H. Wiebe, J.W. Martin, & S.C. France. 1992. Deep-sea amphipod swarms. Nature, 358: 25-26 (

France, S. C., R. C. Vrijenhoek & R. R. Hessler. 1992. Genetic differentiation between spatially-disjunct populations of the deep-sea, hydrothermal vent-endemic amphipod Ventiella sulfuris. Marine Biology 114: 551-559[ABSTRACT] (

Brusca, R. C. & S. C. France. 1992. The genus Rocinela (Crustacea: Isopoda: Aegidae) in the tropical eastern Pacific. Zoological Journal of the Linnean Society 106: 231-275

Hessler, R. R., S. C. France, M. A. Boudrias. 1987. Hydrothermal Vent Communities of the Marianas Back-Arc Basin. Eos 68(44): 1531 (Abstract)

Popular literature:

Pante, E. & S.C. France. 2009. Randonnée au fond des océans: l'exploration des monts sous-marins. Le Courrier de la Nature 247: 34-41

France, S.C. 2004. "Anthozoa." In: Schlager, N., ed., Grzimek's Animal Life Encyclopedia, Vol. 1: Lower Metazoans and Lesser Deuterostomes, Schlager Group Inc., Chicago.

Hessler, R. R., S. C. France & M. A. Boudrias. 1991. "The Ocean Depths." In: Encyclopedia of the Earth: Oceans and Islands, F. H. Talbot and R. E. Stevenson, eds., Smithmark, New York.

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